Researches on Irritability of Plants

Arrest at diastole by the male of anode, and systolic contraction by the break of anode . . Response of Biophytum under increasing intensity of stimulus Multiple response of Biophytum Drie a Sigele Cree electric shock . Multiple response in Avervhoa : : : 4 Multiple response under strong thermal shock . : Multiple response under strong chemical stimulation 146. Gradual diminution of amplitude under isolation . 4 35S 147. Stoppage of pulsation brought on by depletion of energy 314 148. Single response to moderate induction shock, in leaflet at standstill < é - : - «14-308 149. Multiple excitation under tetanisation . - 308 150. Single response to stimulus of light of brief ‘duratiee 310 151. Restoration of pulsation under continued action of light 307

152. Arrest of pulsation by. ligature . : : + epee 153. Arrest of pulsation by the action of a cut . = c Lsow 154. Determination of apex time - : - 5 Poet tc0) 155. Refractory period of Desmodium . - : : Benne io) 156. Incapability of tetanus < = 7315 157. Extra pulsation induced by se hs of ere see lation during diastole. - 359 158. Inhibitory effect of transmitted Seetion on rhythmic f pulsation oe 0 159. Augmentation of pulsation induced” by ‘transmitted excitation : - - 2 - - = omg

160. Effect of lowering of temperature : 2 : = 325 161. Effect of rapid cooling . : : -meSeO 162. Arrest of pulsation at low temperature - - 326 163. Effect of rise of temperature on amplitude and period 2 27, 164. Effect of continuous rise of temperature : Pemee, 1) 165. Effect of continuous rise and return to original tem- perature : eae 166. Effect of variation of temperature « on eyctelic and diastolic 178. Quick arrest of pulsation by the action of KCN platen 339

179. Arrest of pulsation at diastole by the action of dilute lactic acid - 340 Acrp and alkali, antagonistic action of, on cardiac pulsation, 339 Additive effect, 54 3 on quantitative relation of, 55 Age, effect of, on polar excitation, 267 Alcohol, effect of, on Mimosa response, 91 a » 9, Desmodium pulsation, 333 ? All-or-none ’ principle, in response of Biophytum, 281 rr ass 3 », cardiac tissue, 282 Ammonia, effect of, on Mimosa response, 94 Anesthetics, action of, on conductivity, 154 », response of Mimosa, 92

» make effect of, on Desmodium pulsation, 236 Apex time, determination of, in Mimosa, 37 » 9», Desmodium, 309 Ascending and descending currents, time difference in the actions of, 204 Pp electrotonic effects of, 165, 239 Ascending and “descending induction shock, time difference in the actions of, 206 Automatic exciter, 66 Automatic response, continuity with multiple, 288 due to storage of energy, 287 cessation of, by depletion of energy, 312 see also Desmodium pulsation Autonomous response, see Automatic response Averrhoa bilimbi, polar action of current on, 228 Averrhoa carambola, multiple response in, 286 5 #, polar action of current in, 226, 262 * \s velocity of transmission of excitation in, 151

Biophytum sensitivum, arrest of excitatory impulse in, by poison, 169 s 5S comparative sensitiveness of animal and, 249 Be 5 direction of preferential conduction in, 152 5 &S dual impulse in, 181 Biophytum sensitivum, effect of increasing intensity of stimulus on, 284 * 53 latent period of, 281 CARBON disulphide, effect of, on Desmodium pulsation, 338 “n »» 9) 9) Mimosa response, 92 Carbonic acid, effect of, on pulsation of Desmodium, 335 » 9) +99 Lesponse in Mimosa, 91 Cardiac pulsation, similarity between Desmodium pulsation and, 321, 323, Chemical agents, effect of, on rhythmic response, 332 »» 9) 99 Mechanical response of Mimosa, 85 Chloroform, effect of, on Desmodium pulsation, 336 » response of Mimosa, 93 Coal '2aS, effect of, on Mimosa response, 92 Coercer, 15 Cold, arrest of Desmodium pulsation by, 326 ,, diminished velocity of transmission by, 162 », excitatory action by sudden application of, 100 5, paralysis of conductivity by, 163 ,, prolongation of latent period by, 130 unmasking of positive impulse by, 195 Condenser discharge, stimulation by, 27 Conduction of excitation, channels of, 133 of a determination of velocity of, 136 5 * effect of fatigue on velocity of, 147 = 5s » ,, intensity of stimulus on velocity of, 142 “4 . » 4, Optimum condition on velocity of, 145 % ms » 4, temperature on velocity of, 148 a ,, 4, preferential direction of, 151 i velocity of, in Averryhoa, 151 . A os Biophytum, 15 x », Mimosa, 141 Constant current, polar effect of, 198 Contact, electrolytic, 68

Death-point, translocation of, by fatigue, 106 ”, ” ” ”? poison, 106 Death-spasm in plants, 98 Desmodium, extra pulsation by induction shock, 319 $5 response of terminal leaflet of, 45 Desmodium pulsation, continuous record of, 294 . determination of apex time, 309 », latent period, 308 ,, refractory period, 310 effect of acid on, 340 alcohol on, 334 alkali on, 340 anesthetics on, 336 oe x » », anode and kathode break on, 237 anode and kathode make on, 237 carbon disulphide on, 338

” Fr 5» », imternal hydrostatic pressure on, 300 =f 3 op yp IRISH UES CoNa, SYoy BS 5 » », Sugar solution on, 333 = - » », transmitted excitation on, 320 $3 > gradual stoppage of, by depletion of energy, 313 on 3 incapability of tetanus, 318 an es measurement of amplitude and period of, 296 3 . similarity between Biophytum pulsation and, 288 - is * 3 cardiac pulsation and, 317, 323, a 5 temperature maximum in, 330 cs a3 a minimum in, 326 3 time relations of, 297 Desmodium at standstill, by cut, 303 “A 3) 5 depletion of energy, 313 ” ” ” ligature, 302 - " 5, revival of pulsation in, by electric shock, 308 ” ” ” ” ” ” stimulus of light, 315 Differential excitability, existence of, in Mimosa pulvinus, 45 oF, method, velocity of transmission by, 138 Direct and indirect ‘effect of stimulus, laws of, 196 Y) 35 <5 2 exhibition of, under different

stimuli, 179 Dual impulses in Averrhoa cavambola, 184 x 3 » Biophytum sensitivum, 180 rs » multiple excitation induced in Biophytum by, 249 » response, detection of excitation by, 98 Es multiple, 283 Bless teen stimulator, 25 Electro-tonic block, arrest of excitatory impulse by, 164, 238 Electrotonus, modification of excitability by, 164 Energy, effect of run-down of, on rhythmic pulsation, 313 », storage of, from external stimulus, 287 Ether, effect of, on Desmodium pulsation, 336

a », Mimosa, 92 Excitability, diurnal variation of, 46 5 effect of temperature on, 60 a electro-tonic variation of, 238 > variation of, by chemical agents, 85 Excitatory impulse, arrest of, by action of poison, 167 = AS 3 5) (COld tO oe reversal, under continuous stimulation in plant and animal, 81 Friction, errors in record introduced by, 9 HABERLANDT on theory of mechanical transmission, 155 Hydro-mechanical theory, 154 Hydrostatic pressure, effect on Desmodium pulsation, 300

INTERMITTENT contact, advantage of, in record, 21 Inversion, thermo-mechanical, 104 KATHODE, depression of excitability at break of, 242 y diastolic expansion by break of, 237 p excitatory effect of, at make, 201 a reduction of diastolic expansion in Desmodium by make of, 236 y, systolic arrest by make of, 237 LATENT period, apparatus for determination of, 111 determination of, in Biophytum, 280 ,, Mimosa, 120 os “4 » » Neptunia, 120 effect of ‘fatigue on, 128 optimum condition on, 128 strength of stimulus on, 125 » 9», temperature on, 130 Taw: of Gimect and indirect effects of sepsice. 196 »» ») polar action of current on plant tissues, 233, 265 Ligature, effect of, on Desmodium pulsation, 303 5 Stannius’, on cardiac pulsation, 302 Light, initiation of pulsatory movement in Desmodium by the action of, 315 Light, multiple response induced by, in Biophytum, 286 Load, effect of, on mechanical response, 56 35 a » ») amount of work performed, 58

Multiple response to strong stimulus in Biophytum, 284 under chemical stimulus, 286 constant current, 249 Neptunia oleracea, latent period of, 120 x ») polar effects of current on, 225 “3 response of, 43 Nitrogen dioxide, abolition of response by the action of, 95 PeEriopic starter, 66 Pfeffer on hydro-mechanical theory, 155 Pfliiger on law of polar excitation in animal tissue, 232, 276 Plant, on relative sensitiveness of animal and, 249 Poison, block of conduction of excitation by local application of, 167 », abolition of pulsation in Desmodium by, 338

Polar action of currents, effect of tissue modification on, 266 RECORD, fixation of, 18 Recorder, optical lever, 178 V4 oscillating, 279 ep resonant, 14 Response, different types of, 64 7 effect of fatigue on, 72 PA io ,, intensity of stimulus on, 61 ,, optimum condition on, 63 Response, electrical, 98, 177 positive and negative, 177, 181 a relation of stimulus to, 62 Rhythmic tissue, similarity of response in animal ana plant, 321 SENSITIVENESS, comparison between animal and plant, 249 ‘Sensitive’ plant, arbitrary distinction between ordinary and, 44 Staircase response in Mimosa, 77 Stannius’ ligature on cardiac pulsation, 302 Stimulation, automatic, 66 chemical method of, 23 electrical methods of, 27, 28, 30

5% thermal method of, 25 Stimulus, direct effect of, 195 indirect effect of, 196 relation between response and, 61 Sugar solution, stimulating action of, on Desmodium pulsation, 333 Sulphur dioxide, effect of, on response of Mimesa, 96 Sulphuretted hydrogen, toxic action of, 94 TEMPERATURE, arrest of Desmodium pulsation by lowering of, 326 a S ,, transmission of excitation by lowering of, 161 effect of, on cardiac pulsation, 323 », Desmodium pulsation, 325 », excitatory efficiency of induction shock, 246 polar action of current, 247

. »» 9» 9) velocity of transmissiun of excitation, 148 maximum for Desmodium pulsation, 331 VELocitTy of excitatory impulse, determination of, by direct method, 135 ” ” ” ” as differential method, 138 5 op a effect of cold on, 161 Work, effect of load on, in Mimosa, 56 measurement of, in Mimosa, 57 rate of, in Mimosa, 58